Shao, G.-J., Sork, V., Wang, Y.-L., Li, L., Chen, H.-F., Wang, W.-J., Huang, M.-Y., Li, Q., and Zhao, J.-L. (2023). Selfing species has greater genetic diver- sity and less structure than related outcrossing species due to seed dispersal and population history in roscoea (zingiberaceae).

Li, L., Zhang, J., Sork, V., Fitzpatrick, M., Liao, H., Wang, G., Sun, Y.-B., Li, Q., and Zhao, J.-L. (2023). Local adaptation-induced evolutionary trap in alpine plants under climate change.

Leonard, J. A., von Holdt, B., Smith, T. B., Sork, V. L., Shapiro, B., Ostrander, A., Rieseberg, L. H., Van Valkenburgh, B., and Lohmueller, K. E. (2023). Remembering distinguished professor robert k. wayne. Molecular ecology, 32(9):2364–2368.

Fitz-Gibbon, S., Mead, A., O’Donnell, S., Li, Z.-Z., Escalona, M., Beraut, E., Sacco, S., Marimuthu, M. P., Nguyen, O., and Sork, V. L. (2023). Reference genome of california walnut, juglans californica, and resemblance with other genomes in the order fagales. Journal of Heredity, page esad036.


Sork, V. L., Cokus, S. J., Fitz-Gibbon, S. T., Zimin, A. V., Puiu, D., Garcia, A., Gugger, P. F., Henriquez, C. L., Zhen, Y., Lohmueller, K. E., et al. (2022). High-quality genome and methylomes illustrate features underlying evolutionary success of oaks. Nature communications, 13(1):2047.

Shaffer, H. B., Toffelmier, E., Corbett-Detig, R. B., Escalona, M., Erickson, B., Fiedler, P., Gold, M., Harrigan, R. J., Hodges, S., Luckau, T. K., et al. (2022). Landscape genomics to enable conservation actions: the california conservation genomics project. Journal of Heredity, 113(6):577–588.

Li, Y., Zhang, X., Wang, L., Sork, V. L., Mao, L., and Fang, Y. (2022). Influence of pliocene and pleistocene climates on hybridization patterns between two closely related oak species in china. Annals of Botany, 129(2):231–245.

Hvilsom, C., Segelbacher, G., Ekblom, R., Fischer, M. C., Laikre, L., Leus, K., O’Brien, D., Shaw, R., and Sork, V. (2022). Selecting species and populations for monitoring of genetic diversity. IUCN Publication.

Fermin, M. M. E., Green, R. E., Toffelmier, E., Corbett-Detig, R., Erickson, B., Fiedler, P., Gold, M., Harrigan, R., Hodges, S., Luckau, T., et al. (2022). Can genomics help save california’s biodiversity? the california conservation genomics project. In Plant and Animal Genome XXIX Conference (January 8-12, 2022). PAG.

Wright, J. W., Ivey, C. T., Canning, C., and Sork, V. L. (2021). Timing of bud burst is associated with climate of maternal origin in quercus lobata progeny in a common garden. Madron˜o, 68(4):443–449.


Sork, V. L. and Wright, J. W. (2021). Replanting oaks? new research in valley oak may help inform seed-sourcing decisions. Protecting California’s native flora since 1965, page 13.

O’Donnell, S. T., Fitz-Gibbon, S. T., and Sork, V. L. (2021). Ancient intro- gression between distantly related white oaks (quercus sect. quercus) shows evidence of climate-associated asymmetric gene exchange. Journal of Hered- ity, 112(7):663–670.

Marden, E., Abbott, R. J., Austerlitz, F., Ortiz-Barrientos, D., Baucom, R. S., Bongaerts, P., Bonin, A., Bonneaud, C., Browne, L., Alex Buerkle, C., et al. (2021). Sharing and reporting benefits from biodiversity research.

Gugger, P. F., Fitz-Gibbon, S. T., Albarr´an-Lara, A., Wright, J. W., and Sork, V. L. (2021). Landscape genomics of quercus lobata reveals genes involved in local climate adaptation at multiple spatial scales. Molecular Ecology, 30(2):406–423.

Browne, L., MacDonald, B., Fitz-Gibbon, S., Wright, J. W., and Sork, V. L. (2021). Genome-wide variation in dna methylation predicts variation in leaf traits in an ecosystem-foundational oak species. Forests, 12(5):569.


Ramirez, H.-L., Ivey, C. T., Wright, J. W., MacDonald, B. W., and Sork, V. L. (2020). Variation in leaf shape in a quercus lobata common garden: tests for adaptation to climate and physiological consequences. Madron˜o, 67(2):77–84.

O’Leary, E. S., Shapiro, C., Toma, S., Sayson, H. W., Levis-Fitzgerald, M., Johnson, T., and Sork, V. L. (2020). Creating inclusive classrooms by engag- ing stem faculty in culturally responsive teaching workshops. International Journal of STEM education, 7:1–15.

Medeiros, C. D. B., Sorfazian, A., Mead, A., Sork, V. L., and Sack, L. (2020). Evolution of drought tolerance across california native oaks. In 2020 ESA Annual Meeting (August 3-6). ESA.

Hipp, A. L., Manos, P. S., Hahn, M., Avishai, M., Bod´en`es, C., Cavender-Bares, J., Crowl, A. A., Deng, M., Denk, T., Fitz-Gibbon, S., et al. (2020). Genomic landscape of the global oak phylogeny. New Phytologist, 226(4):1198–1212.

Dias Barros Medeiros, C., Sorfazian, A., Mead, A., Sork, V., and Sack, L. (2020). Evolution of drought tolerance across california native oaks. In AGU Fall Meeting Abstracts, volume 2020, pages B082–0014.

Browne, L., Mead, A., Horn, C., Chang, K., A. Celikkol, Z., L. Henriquez, C., Ma, F., Beraut, E., S. Meyer, R., and Sork, V. L. (2020). Experimental dna demethylation associates with changes in growth and gene expression of oak tree seedlings. G3: Genes, Genomes, Genetics, 10(3):1019–1028.

Older publications (2020-2010)

A full publications list pre-2010 is available on Google Scholar

Zhao, J.-L., Werth, S., Ikegami, M., Gugger, P. F., and Sork, V. L. (2019). Historical interactions are predicted to be disrupted under future climate change: The case of lace lichen and valley oak. Journal of Biogeography, 46(1):19–29.

Mead, A., Penaloza Ramirez, J., Bartlett, M. K., Wright, J. W., Sack, L., and Sork, V. L. (2019). Seedling response to water stress in valley oak (quercus lobata) is shaped by different gene networks across populations. Molecular Ecology, 28(24):5248–5264.

Burge, D. O., Parker, V. T., Mulligan, M., and Sork, V. L. (2019). Influence of a climatic gradient on genetic exchange between two oak species. American Journal of Botany, 106(6):864–878.

Browne, L., Wright, J. W., Fitz-Gibbon, S., Gugger, P. F., and Sork, V. L. (2019). Adaptational lag to temperature in valley oak (quercus lobata) can be mitigated by genome-informed assisted gene flow. Proceedings of the National Academy of Sciences, 116(50):25179–25185.

Wanderley, A. M., Machado, I. C. S., de Almeida, E. M., Felix, L. P., Galetto, L., Benko-Iseppon, A. M., and Sork, V. L. (2018). The roles of geography and environment in divergence within and between two closely related plant species inhabiting an island-like habitat. Journal of Biogeography, 45(2):381– 393.

Sork, V. L. (2018). Genomic studies of local adaptation in natural plant popu- lations. Journal of Heredity, 109(1):3–15.

Ortego, J., Gugger, P. F., and Sork, V. L. (2018). Genomic data reveal cryptic lineage diversification and introgression in californian golden cup oaks (section protobalanus). New Phytologist, 218(2):804–818.

Oney-Birol, S., Fitz-Gibbon, S., Chen, J.-M., Gugger, P. F., and Sork, V. L. (2018). Assessment of shared alleles in drought-associated candidate genes among southern california white oak species (quercus sect. quercus). BMC genetics, 19:1–12.

Martins, K., Gugger, P. F., Llanderal-Mendoza, J., Gonz´alez-Rodr´ıguez, A., Fitz-Gibbon, S. T., Zhao, J.-L., Rodr´ıguez-Correa, H., Oyama, K., and Sork, V. L. (2018). Landscape genomics provides evidence of climate-associated genetic variation in mexican populations of quercus rugosa. Evolutionary Applications, 11(10):1842–1858.

Kim, B. Y., Wei, X., Fitz-Gibbon, S., Lohmueller, K. E., Ortego, J., Gugger, P. F., and Sork, V. L. (2018). Radseq data reveal ancient, but not pervasive, introgression between californian tree and scrub oak species (quercus sect. quercus: Fagaceae). Molecular Ecology, 27(22):4556–4571.

Gugger, P. F., Liang, C. T., Sork, V. L., Hodgskiss, P., and Wright, J. W. (2018). Applying landscape genomic tools to forest management and restora- tion of hawaiian koa (acacia koa) in a changing environment. Evolutionary Applications, 11(2):231–242.

Cannon, C. H., Brendel, O., Deng, M., Hipp, A. L., Kremer, A., Kua, C.- S., Plomion, C., Romero-Severson, J., and Sork, V. L. (2018). Gaining a global perspective on fagaceae genomic diversification and adaptation. New Phytologist, 218(3):894–897.

Browne, L., Wright, J. W., Gugger, P., and Sork, V. L. (2018b). Integrating genomic data and common garden experiments to predict the growth of valley oak (quercus lobata) in a changing climate. In 2018 ESA Annual Meeting (August 5–10). ESA.

Browne, L., Ottewell, K., Sork, V. L., and Karubian, J. (2018a). The relative contributions of seed and pollen dispersal to gene flow and genetic diversity in seedlings of a tropical palm. Molecular ecology, 27(15):3159–3173.

Peguero, G., Bonal, R., Sol, D., Mun˜oz, A., Sork, V. L., and Espelta, J. M. (2017). Tropical insect diversity: evidence of greater host specialization in seed-feeding weevils. Ecology, 98(8):2180–2190.

Ortego, J., Gugger, P. F., and Sork, V. L. (2017). Impacts of human-induced environmental disturbances on hybridization between two ecologically differ- entiated californian oak species. New Phytologist, 213(2):942–955.

Gugger, P. F., Pen˜aloza-Ram´ırez, J. M., Wright, J. W., and Sork, V. L. (2017). Whole-transcriptome response to water stress in a california endemic oak, quercus lobata. Tree physiology, 37(5):632–644.

Gienapp, P., Fior, S., Guillaume, F., Lasky, J. R., Sork, V. L., and Csill´ery, K. (2017). Genomic quantitative genetics to study evolution in the wild. Trends in Ecology & Evolution, 32(12):897–908.

Fitz-Gibbon, S., Hipp, A. L., Pham, K. K., Manos, P. S., and Sork, V. L. (2017). Phylogenomic inferences from reference-mapped and de novo assembled short- read sequence data using radseq sequencing of california white oaks (quercus section quercus). Genome, 60(9):743–755.

Wright, J. W. and Sork, V. L. (2016). Early results from a newly-established provenance test in valley oak (quercus lobata) show significant population differentiation. In Proceedings of Workshop on Gene Conservation of Tree Species–Banking on the Future, page 164.

Wei, X., Sork, V. L., Meng, H., and Jiang, M. (2016). Genetic evidence for central-marginal hypothesis in a cenozoic relict tree species across its distri- bution in china. Journal of Biogeography, 43(11):2173–2185.

Verhoeven, K. J., Vonholdt, B. M., and Sork, V. L. (2016). Epigenetics in ecology and evolution: what we know and what we need to know.

Sork, V. L., Squire, K., Gugger, P. F., Steele, S. E., Levy, E. D., and Eckert, J. (2016d). Landscape genomic analysis of candidate genes for climate adaptation in a california endemic oak, quercus lobata. American journal of botany, 103(1):33–46.

Sork, V. L., Riordan, E., Gugger, P. F., Fitz-Gibbon, S., Wei, X., and Ortego, J. (2016c). Phylogeny and introgression of california scrub white oaks (quercus section quercus). International Oaks, 27:61–74.

Sork, V. L., Gugger, P. F., Chen, J.-M., and Werth, S. (2016b). Evolutionary lessons from california plant phylogeography. Proceedings of the National Academy of Sciences, 113(29):8064–8071.

Sork, V. L., Fitz-Gibbon, S. T., Puiu, D., Crepeau, M., Gugger, P. F., Sherman, R., Stevens, K., Langley, C. H., Pellegrini, M., and Salzberg, S. L. (2016a). First draft assembly and annotation of the genome of a california endemic oak quercus lobata n´ee (fagaceae). G3: Genes, Genomes, Genetics, 6(11):3485– 3495.

Sork, V. L. (2016). Gene flow and natural selection shape spatial patterns of genes in tree populations: implications for evolutionary processes and appli- cations. Evolutionary applications, 9(1):291–310.

Riordan, E. C., Gugger, P. F., Ortego, J., Smith, C., Gaddis, K., Thompson, P., and Sork, V. L. (2016). Association of genetic and phenotypic variability with geography and climate in three southern california oaks. American journal of botany, 103(1):73–85.

Gugger, P. F., Fitz-Gibbon, S., PellEgrini, M., and Sork, V. L. (2016b). Species- wide patterns of dna methylation variation in quercus lobata and their asso- ciation with climate gradients. Molecular Ecology, 25(8):1665–1680.

Gugger, P. F., Cokus, S. J., and Sork, V. L. (2016a). Association of transcriptome-wide sequence variation with climate gradients in valley oak (quercus lobata). Tree Genetics & Genomes, 12:1–14.

Gaddis, K. D., Thompson, P. G., and Sork, V. L. (2016). Dry-washes determine gene flow and genetic diversity in a common desert shrub. Landscape Ecology, 31:2215–2229.

Chen, J.-M., Werth, S., and Sork, V. L. (2016). Comparison of phylogeograph- ical structures of a lichen-forming fungus and its green algal photobiont in western north america. Journal of Biogeography, 43(5):932–943.

Bonal, R., Espelta, J. M., Munoz, A., Ortego, J., Aparicio, J. M., Gaddis, K., and Sork, V. L. (2016). Diversity in insect seed parasite guilds at large geographical scale: the roles of host specificity and spatial distance. Journal of biogeography, 43(8):1620–1630.

Sork, V. L., Smouse, P. E., Grivet, D., and Scofield, D. G. (2015). Impact of asymmetric male and female gamete dispersal on allelic diversity and spatial genetic structure in valley oak (quercus lobata n´ee). Evolutionary ecology, 29:927–945.

Platt, A., Gugger, P. F., Pellegrini, M., and Sork, V. L. (2015). Genome-wide signature of local adaptation linked to variable c p g methylation in oak populations. Molecular Ecology, 24(15):3823–3830.

Ortego, J., Noguerales, V., Gugger, P. F., and Sork, V. L. (2015b). Evolutionary and demographic history of the californian scrub white oak species complex: an integrative approach. Molecular Ecology, 24(24):6188–6208.

Ortego, J., Gugger, P. F., and Sork, V. L. (2015a). Climatically stable land- scapes predict patterns of genetic structure and admixture in the californian canyon live oak. Journal of Biogeography, 42(2):328–338.

Cokus, S. J., Gugger, P. F., and Sork, V. L. (2015). Evolutionary insights from de novo transcriptome assembly and snp discovery in california white oaks. BMC genomics, 16(1):1–19.

Albarr´an-Lara, A. L., Wright, J. W., Gugger, P. F., Delfino-Mix, A., Pen˜aloza- Ram´ırez, J. M., Sork, V. L., et al. (2015). Phenotypic variation in california populations of valley oak (quercus lobata n´ee) sampled along elevational gra- dients. In Proceedings of the seventh oak symposium: managing oak woodlands in a dynamic world: Vol. General Technical Report PSW-GTR, volume 251.

Werth, S. and Sork, V. L. (2014). Ecological specialization in trebouxia (tre- bouxiophyceae) photobionts of ramalina menziesii (ramalinaceae) across six range-covering ecoregions of western north america. American journal of botany, 101(7):1127–1140.

Thompson, P. G., Smouse, P. E., Scofield, D. G., and Sork, V. L. (2014). What seeds tell us about birds: a multi-year analysis of acorn woodpecker foraging movements. Movement ecology, 2(1):1–10.

Sork, V. L. and Werth, S. (2014). Phylogeography of r amalina menziesii, a widely distributed lichen-forming fungus in western n orth a merica. Molecular Ecology, 23(9):2326–2339.

Ortego, J., Gugger, P. F., Riordan, E. C., and Sork, V. L. (2014). Influence of climatic niche suitability and geographical overlap on hybridization patterns among southern californian oaks. Journal of Biogeography, 41(10):1895–1908.

Mix, A. D., Wright, J. W., Gugger, P. F., Liang, C., Sork, V. L., et al. (2014). Establishing a range-wide provenance test in valley oak (quercus lobata n´ee) at two california sites. In California Oak Symposium: Managing Oak Wood- lands in a Dynamic World, page 413.

Gaddis, K. D., Zukin, H. L., Dieterich, I. A., Braker, E., and Sork, V. L. (2014b). Effect of clonal reproduction on genetic structure in pentaclethra macroloba (fabaceae: Mimosoideae). Revista de Biolog´ıa Tropical, 62(2):443–454.

Gaddis, K. D., Zukin, H. L., Dieterich, I. A., Braker, E., and Sork, V. L. (2014a). Efecto de la reproducci´on clonal en la estructura gen´etica de pentaclethra macroloba (fabaceae: Mimosoideae). Revista de Biolog´ıa Tropical, 62(2):443– 454.

Aguilar-Barajas, E., Sork, V. L., Gonz´alez-Zamora, A., Rocha-Ram´ırez, V., Arroyo-Rodr´ıguez, V., and Oyama, K. (2014). Isolation and characteriza- tion of polymorphic microsatellite loci in spondias radlkoferi (anacardiaceae). Applications in Plant Sciences, 2(11):1400079.

Sork, V., Aitken, S., Dyer, R., Eckert, A., Legendre, P., and Neale, D. (2013). Putting the landscape into the genomics of trees: approaches for understand- ing local adaptation and population responses to changing climate. Tree Genetics & Genomes, 9:901–911.

Rose, A., Bernatchez, L., Aur´elie, B., Buerkle, C. A., Emerson, B. C., Garant, D., Giraud, T., Nolan, K., Sean, R., Slate, J., et al. (2013). A road map for molecular ecology.

Petit, R. J., Carlson, J., Curtu, A. L., Loustau, M.-L., Plomion, C., Gonz´alez- Rodr´ıguez, A., Sork, V., and Ducousso, A. (2013). Fagaceae trees as models to integrate ecology, evolution and genomics. New Phytologist, 197(2):369–371.

Moroney, J. R., Rundel, P. W., and Sork, V. L. (2013). Phenotypic plastic- ity and differentiation in fitness-related traits in invasive populations of the mediterranean forb centaurea melitensis (asteraceae). American journal of botany, 100(10):2040–2051.

Herrera-Arroyo, M. L., Sork, V. L., Gonz´alez-Rodr´ıguez, A., Rocha-Ram´ırez, V., Vega, E., and Oyama, K. (2013). Seed-mediated connectivity among frag- mented populations of quercus castanea (fagaceae) in a mexican landscape. American Journal of Botany, 100(8):1663–1671.

Gugger, P. F., Ikegami, M., and Sork, V. L. (2013). Influence of late q uaternary climate change on present patterns of genetic variation in valley oak, q uercus lobata n´ee. Molecular Ecology, 22(13):3598–3612.

Andrew, R. L., Bernatchez, L., Bonin, A., Buerkle, C. A., Carstens, B. C., Emerson, B. C., Garant, D., Giraud, T., Kane, N. C., Rogers, S. M., et al. (2013). A road map for molecular ecology. Molecular ecology, 22(10):2605– 2626.

Smouse, P. E., Sork, V. L., Scofield, D. G., and Grivet, D. (2012). Using seedling and pericarp tissues to determine maternal parentage of dispersed valley oak recruits. Journal of Heredity, 103(2):250–259.

Scofield, D. G., Smouse, P. E., Karubian, J., and Sork, V. L. (2012). Use of alpha, beta, and gamma diversity measures to characterize seed dispersal by animals. The American Naturalist, 180(6):719–732.

Robledo-Arnuncio, J. J., Grivet, D., Smouse, P., and Sork, V. (2012). Psa: software for parental structure analysis of seed or seedling patches. Molecular Ecology Resources, 12(6):1180–1189.

Ortego, J., Riordan, E. C., Gugger, P. F., and Sork, V. L. (2012). Influence of environmental heterogeneity on genetic diversity and structure in an endemic southern californian oak. Molecular Ecology, 21(13):3210–3223.

Gonz´alez-Zamora, A., Arroyo-Rodrıguez, V., Oyama, K., Sork, V., Chapman, C., et al. (2012). Sleeping sites and latrines of spider monkeys in continuous and. SITIOS DORMIDERO Y LETRINAS DE MONOS ARAN˜A (Ateles ge- offroyi) EN BOSQUE CONTINUO Y FRAGMENTADO: IMPLICACIONES PARA LA DISPERSIO´N DE SEMILLAS, page 35.

Gaddis, K. D. and Sork, V. L. (2012). Connectivity of a patchy mojave desert shrub: Examining landscape genetics at multiple spatial and temporal scales. In 97th ESA Annual Meeting.

Scofield, D. G., Alfaro, V. R., Sork, V. L., Grivet, D., Martinez, E., Papp, J., Pluess, A. R., Koenig, W. D., and Smouse, P. E. (2011). Foraging patterns of acorn woodpeckers (melanerpes formicivorus) on valley oak (quercus lobata n´ee) in two california oak savanna-woodlands. Oecologia, 166:187–196.

S´anchez-Humanes, B., Sork, V. L., and Espelta, J. M. (2011). Trade-offs be- tween vegetative growth and acorn production in quercus lobata during a mast year: the relevance of crop size and hierarchical level within the canopy. Oecologia, 166:101–110.

Rosas, F., Quesada, M., Lobo, J. A., and Sork, V. L. (2011). Effects of habitat fragmentation on pollen flow and genetic diversity of the endangered tropical tree swietenia humilis (meliaceae). Biological Conservation, 144(12):3082– 3088.

Werth, S. and Sork, V. L. (2010). Identity and genetic structure of the pho- tobiont of the epiphytic lichen ramalina menziesii on three oak species in southern california. American Journal of Botany, 97(5):821–830.

Wang, H., Sork, V. L., Wu, J., and Ge, J. (2010). Effect of patch size and isolation on mating patterns and seed production in an urban population of chinese pine (pinus tabulaeformis carr.). Forest Ecology and Management, 260(6):965–974.

Waits, L., Sork, V., et al. (2010). Landscape genetics. Molecular Ecology, 19(17):3489–3835.

Sork, V. L., Davis, F. W., Westfall, R., Flint, A., Ikegami, M., Wang, H., and Grivet, D. (2010). Gene movement and genetic association with regional climate gradients in california valley oak (quercus lobata n´ee) in the face of climate change. Molecular ecology, 19(17):3806–3823.

Scofield, D. G., Sork, V. L., and Smouse, P. E. (2010). Influence of acorn woodpecker social behaviour on transport of coast live oak (quercus agrifolia) acorns in a southern california oak savanna. Journal of Ecology, 98(3):561– 572.

Karubian, J., Sork, V. L., Roorda, T., Dur˜aes, R., and Smith, T. B. (2010). Destination-based seed dispersal homogenizes genetic structure of a tropical palm. Molecular Ecology, 19(8):1745–1753.